Words Containing: L,E,A,N,I,N,G
(In Any Order)
There are 1,136 words,
2,159 phrases and
0 abbr's with
L,E,A,N,I,N,G in.
Best Scoring Words With: L,E,A,N,I,N,G
More words | |||||||
---|---|---|---|---|---|---|---|
Rank | Word | Save? | Length | Usage | Points | Type | |
1 | penalizing | 10 | 22 | verb, adjectivev, adj | |||
2 | melanizing | 10 | 22 | verb, adjectivev, adj | |||
3 | javelining | 10 | 21 | verbv | |||
4 | explaining | 10 | 20 | verbv | |||
5 | explanting | 10 | 20 | verbv | |||
6 | blackening | 10 | 19 | verb, noun, adjectivev, n, adj | |||
7 | necklacing | 10 | 19 | verb, noun, adjectivev, n, adj | |||
8 | menacingly | 10 | 18 | adverb, adjectiveadv, adj | |||
9 | sneakingly | 10 | 18 | adverbadv | |||
10 | flankering | 10 | 18 | ||||
or scroll down to see all results... | |||||||
Tip: Scrabble EU allows far more words than US! Also change the max length to see more words. |
Words (58)
alignment12cleansing12leningradcanceling14lamenting12galantine10enslaving13congenial12enlarging11panelling12inelegant10deplaning13meningeal12leavening13bengaline12annealing10fenagling14yeanlings13levanting13enplaning12unsealing10panelings12enclaving15renailing10emplaning14reloaning10weanlings13unaligned11meaningly15learnings10lilangeni10relanding11entailing10eglantine10gantlines10
...View all with 9 letters...
Phrases (3)
conga linein generalangle ironWords (107)
meaningful16englishmanlaundering12fingernail14concealing15intangible13congenital13channeling16unraveling14cancelling15headlining15alienating11changeling17menacingly18slackening17penalizing22tangential11blanketing17repaneling13enfilading15unlearning11nonaligned12elongating12blarneying16blackening19elongation11relearning11unpleasing13engagingly16yearningly17sanderling12ensilaging12congealing14ensnarling11melanizing22
...View all with 10 letters...
Phrases (11)
genus inulaglans penisian flemingonion bagelbengal kinoagonic linedanger lineleading mangiant elandgolden rainlanding netWords (163)
meaningless14challenging18nightingale16eliminating14alternating12channelling17freelancing17enthralling15unravelling15rebalancing16realignment14disentangle13unceasingly17legionnaire12unappealing16langoustine12unnavigable17delineating13preplanning16unrevealing15congelation14interlacing14deafeningly19malingering15languidness13uncongenial14benignantly17unfaltering15lastingness12endearingly16gainfulness15banteringly17infrangible17changelings18repanelling14
...View all with 11 letters...
Phrases (43)
line drawingrange animaldry cleaningmale bondinggene linkagegenus laniusleading tonewaiting linehelping handroman legionwarning bellcleaning padbrake lininglanding deckenabling actgenus lyoniaangular veingenus anolismaginot linele gallienneranging polegenus nolinaheaving linedining tablegarden snailpineal glandenglish beanmin languagegeneral lienleading reingambling denlingual veinindian lodgelean againstblanking die
...View all with 11 letters...
Words (201)
increasingly18unimaginable17neutralizing22congeniality18unflattering16stonewalling16englishwomangenerational13hallucinogen18streamlining15generalizing23convalescing20enchantingly21centralizing24unwaveringly22disentangled15misalignment17meaningfully21nauseatingly16congenitally18evangelizing26unprevailing18inexpugnable24palingenesis15magniloquent24ungainliness13entreatingly16tangibleness15underplaying19generalising14recanalizing24interplaying18interlarding14palingenetic17interleaving16
...View all with 12 letters...
Phrases (72)
alec guinnessgrain cleanergenus enceliaaegean islandnesting placecutting anglevigilance mangenital organbedding plantlanding placepanel heatinggasoline tankgenus blarinabelgian congogenus vanillabelgian francgenus nasalismagic lanterngentleman jimlake winnipeghigh-angle gunheating plantradial enginegenus talinumgenus nigellaeccrine glandlanguage unitsalient anglegenus nitellawankel enginelieutenant jggenus haleniastarting linegenus lansiumplacer mining
...View all with 12 letters...
Words (171)
grandchildren21congressional16contemplating20acknowledging25magnificently24overanalyzing29housecleaning19nonnegotiable16nonregulation14gentlemanlike20meaninglessly19externalizing30interpolating16undiagnosable17unchallenging20grandiloquent24threateningly20penetratingly19internalizing23unfalteringly20nonmeaningful19niggardliness16undeviatingly21personalising16personalizing25retranslating14encapsulating18overbalancing21nonallergenic16hallucinogens19antigenically19precancelling20mischanneling21encouragingly20conglutinated17
...View all with 13 letters...
Phrases (116)
genus glossinagenus anguillabelgian endiveline engravingligneous plantgenus nautilusguardian angelgenus magnoliafemale bondingfinal judgmentgondi languageking of englandvena lingualiscleaning clothalgerian dinarkarl menningermental anguishinland passagefurnace liningeating utensilmental testingfiling cabinetplanning framehindi languagegenus lusciniadarning needlegenus angelicagenus gloxiniaitalian regiongenus boltoniainterior anglelunatic fringegenus encyclialaryngeal veinpagan religion
...View all with 13 letters...
Words (142)
unrecognizable28hallucinogenic22intelligentsia15generalization24unrecognisable19conglomeration19generalisation15entertainingly18emotionalizing26meaningfulness20congregational18unhesitatingly21gelatinousness15oleaginousness15intangibleness17grandiloquence27magniloquently29interpellating17overexplaining27denuclearizing27generationally18universalizing27conglomerating20mischannelling22antiallergenic17credentialling18unappetizingly31nonoverlapping22nonideological18concelebrating21intransigently18magniloquences28ungainlinesses15imaginableness19countervailing20
...View all with 14 letters...
Phrases (178)
negeri sembilangarageman's liennoachian delugebelgian griffonwilling and ablehandling chargeheating elementvalencia orangestinking wattlegenus lagenariawintun languagedandelion greengenus laminarialexical meaningmagnetic needlepingpong paddleinfernal regiondanish languagegenus fistulinaholding patternjuglans cinereaboating licensekingdom of nepalluvian languageeva le galliennegenus fringillacleansing agentanglesea islandserengeti plaingenista anglicagenus gallinagonepali languagegenus livistonalarge intestinegenus consolida
...View all with 14 letters...
Words (125)
interchangeable23meaninglessness18conceptualizing31gentlemanliness18anticholinergic23unchangeability26everlastingness19counterclaiming22conglomerations20decriminalizing30monumentalizing29disenchantingly25interrogational16dishearteningly23impregnableness22transliterating16occidentalizing30conceptualising22generalisations16antiegalitarian16generalizations25interphalangeal21understandingly21investigational19ontogenetically21evangelizations28mongrelizations27miscegenational20unimaginatively24denationalizing26hallucinogenics23organochlorines21encephalitogens23reconsolidating19intangibilities18
...View all with 15 letters...
Phrases (194)
italian dressinggenus potentilladissolving agentpunjabi languagegenus gnaphaliumbengali languageaegadean islandsline of reasoningmanner of walkinggenus phalangiumgenus mandrillusgenus chinchillasinhala languagesignaling devicearthurian legendenglish lavenderspanish languagefencing materialgenus leishmaniatraining collegegenus tillandsiarussian languageeugenia unifloragenus flammulinasteering linkageairplane landinglegal instrumentblistering agentnigerian capitalnitrogen balancemichel montaignegenus dipladeniaantifungal agentlearning abilityanglo-saxon deity
...View all with 15 letters...
Words (59)
gastrointestinalinextinguishablydisentanglementsantitechnologiesinfrangibilitieshyperventilatinginextinguishableimaginablenessesvaingloriousnesspleasant-smellingintangiblenessesconsanguineouslyoleaginousnessesphenomenologicalmaladministeringnanotechnologiesregionalizationsgelatinousnessesself-organisationimmunoregulationself-organizationmeaningfulnessesgovernmentalismsgovernmentalistsgovernmentalizedgovernmentalizesintercorrelatinginward-developingreorganizationalmarginocephaliansupercalenderingintergradationalhemagglutinatingovercentralizingnonbiodegradable
...View all with 16 letters...
Phrases (215)
multiengine planeitalian greyhoundold latin languagedefinite integralgenus periplanetatender loving careathletic traininggreater celandineafrasian languagegenus antilocaprazonal pelargoniumgenus cheilanthesmale genital organschool assignmentgeorgian languagethracian languageisland of guernseypharyngeal tonsilalexander flemingmonongahela rivermagnetic flux unitsignal/noise ratiobalinese languagenorwegian lobsterpolishing machinealbanian languagecynoscion regalispicea engelmanniihellenic languageleading indicatornyamwezi languagealton glenn millermind-altering drugvirginal membranemalignant anaemia
...View all with 16 letters...
Words (37)
indistinguishableintergovernmentalintellectualizingunexchangeabilityantitechnologicalicelandic-speakingantireligiousnessreconceptualizingindefatigablenessmultigenerationaltransgenerationalrecontextualizingimmunogeneticallyrefrangiblenessesgentlemanlikenessreindustrializinggentlemanlinessesimmunoregulationsmeaninglessnessesgovernmentalizingdeindustrializingundistinguishableimpregnablenessesintergenerationalultracentrifugingconventionalizinghemagglutinationsprofessionalizingnonbiodegradablesdepartmentalizingcounterblockadinghaemagglutinationunchangeabilitiesscandalmongeringscongregationalism
...View all with 17 letters...
Phrases (195)
castilian languagelandscape paintingnikolaas tinbergenmalaysian languagesupernatural beingjohn the evangelistgenus phalaenopsismaterials handlingyeniseian languageafrican golden wolfeuropean gallinuleabkhasian languagelactogenic hormonesinhalese languagechinookan languageplastering machinedravidian languageanatolian languageclimbing hydrangeamalignant hepatomamalignant melanomamalignant neoplasmnetherlands guianalong-chain moleculelimanda ferrugineapersonal magnetismpersonal organiserliaodong peninsulacongenital anomalycharles dana gibsonferdinand magellancleaning implementdoctrine of analogygranulator machinemichel de montaigne
...View all with 17 letters...
Words (19)
compartmentalizinginternationalizinginterchangeabilityinexpugnablenessescompartmentalisingvaingloriousnessescongregationalismscongregationalistsphenomenologicallynoninterchangeableunpleasant-smellinginternationalisingindustrial-strengthmicroencapsulatingroentgenologicallyovergeneralizationphytohemagglutininself-aggrandisementself-aggrandizementPhrases (186)
hindustani languagenavigational systemaegean civilisationfulminating mercuryon an irregular basisfemale genital organhallucinogenic drugindefinite integraltransmitting aerialalternating currentprograming languagemusical arrangementparliamentary agentdangling participlehablot knight brownemordvinian languagegian lorenzo berninimilitary governmentantagonistic musclelinguistic relationgenetic abnormalityculpable negligenceludwig van beethovenswedish nightingaleallegheny mountainsinflectional endingangle of inclinationanagasta kuehniellabignonia capreolatafukkianese languageinstallation chargespatial arrangementblue ridge mountainscapital of argentinainterquartile range
...View all with 18 letters...
Words (15)
ultracentrifugationparthenogeneticallyencephalomeningitislanguedoc-roussillongentlemanlikenessesdehydrochlorinatingmeningoencephaliticmeningoencephalitisdendrochronologicalinterorganizationalovergeneralizationsindefatigablenessesoverindustrializingphytohemagglutininsinterchangeablenessPhrases (152)
aquilegia canadensisdelusions of grandeurnational debt ceilingmechanical advantagesalicylate poisoninglanguage requirementgenus branchiobdellagreater sunda islandsindo-iranian languagebearing false witnessgerard manley hopkinsinternal jugular veinaccounting principlegenus delphinapterusimmune gamma globulinglycerinated gelatinaustralian aboriginedepartment of englishflorence nightingaleazerbaijani languagedari persian languagemartin luther king daydiversionary landingcontingent liabilityangolan monetary unitcongressional recordadlai ewing stevensonhindoostani languagehydrangea paniculatagentianella amarellaangular accelerationchinese angelica treepin-tailed sandgrouseterrain intelligencehelianthus giganteus
...View all with 19 letters...
Words (7)
magnetofluiddynamicsinterchangeabilitiesreinstitutionalizingdeinstitutionalizingultracentrifugationsroentgenographicallynephroangiosclerosisPhrases (113)
scandinavian languagedwarf flowering almondbyelorussian languagesingle-valued functionplasminogen activatorlubricating substancegenus hamamelidanthumluscinia megarhynchosvirgil garnett thomsonsamuel rawson gardinermilitary intelligenceaustralian bonytonguephylogenetic relationedward estlin cummingsindustrial managementtwo-dimensional figurealgerian monetary uniteastern turki languagetibeto-burman languageinterphalangeal jointbotaurus lentiginosusgeorg philipp telemannwilliam jennings bryanandrew fielding huxleyjohn kenneth galbraithgentiana quinquefoliadwarf golden chinkapinswainsona galegifoliahymenoxys grandifloraomnidirectional rangeamerican sign languagedivision euglenophytacollective bargainingfamily plantaginaceaenegotiable instrument
...View all with 20 letters...
Words (8)
interchangeablenessesneuroendocrinologicalotorhinolaryngologiesmeningoencephalitidesdendrochronologicallyoverintellectualizingantiferromagneticallyindistinguishablenessPhrases (123)
agglutinative languagebulgarian monetary unitanigozanthus manglesiibillie jean moffitt kingstephen william hawkingsurgical contraceptioninterlingual renditionstrategic intelligencematernal-infant bondingmegaptera novaeangliaecynoglossum officinalecentral processing unitmetallurgical engineereugene gladstone o'neillhaitian creole languagedano-norwegian languagehans holbein the youngerknight of the round tablecongenital abnormalitymongolian monetary unitsyngnathus hildebrandisleeping accommodationmichel eyquem montaignecharles francois gounodjohn fitzgerald kennedyjohn greenleaf whittierringgold wilmer lardnercongressional districtprofessional wrestlingstone splitting machinegentianopsis thermalisdevelopmental learningthin-leaved stringybarkamerican virgin islandsfrancis turner palgrave
...View all with 21 letters...
Words (1)
indistinguishabilitiesPhrases (104)
gregorian calendar monthtransient global amnesianavigational instrumentguatemalan monetary unitthousand island dressingsouth dravidian languagehenry engelhard steinwayfloating-point operationthree-dimensional figuresatellite triangulationsamuel pierpoint langleygoing-out-of-business saledisplaying incompetencebirth-control campaignerengraulis encrasicholusnotemigonus crysoleucasprolonged interrogationedward kennedy ellingtonmichelangelo buonarrotidiplomatic negotiationsedwin arlington robinsonlegionella pneumophiliadilatation and curettagebaron olivier of brightonanalog-digital converternegara brunei darussalamwilliam penn adair rogersgreat mendenhall glacierdendrocalamus giganteusgalina sergeevna ulanovajan evangelista purkinjeblack-crowned night heroncharlotte perkins gilmaneverglades national parkgentianopsis holopetala
...View all with 22 letters...
Words (1)
indistinguishablenessesPhrases (71)
gilbert and ellice islandsaugust wilhelm von hofmanncapital of papua new guineabangladeshi monetary unitgiovanni battista tiepolotechnological revolutionchronological successionkingdom of the netherlandstrigonella foenumgraecumlarge indefinite quantitycaprimulgus carolinensisjean louis charles garnieroxford english dictionarylipid-lowering medicationgentianella quinquefolianewton's law of gravitationhelianthus angustifoliusiris germanica florentinaasplenium adiantum-nigrumgreater new orleans bridgemachine-oriented languagefluctuating exchange rategastrointestinal hormonerenewable energy industrycerebrospinal meningitispagophilus groenlandicusjapanese flowering cherryanoplophora glabripennisnongonococcal urethritisbabylonian weeping willowhanging gardens of babylonjudgement on the pleadingshugueninia tanacetifoliaproblem-oriented languagegabriel daniel fahrenheit
...View all with 23 letters...
Words (1)
laryngotracheobronchitisPhrases (69)
carnegie mellon universityprofessional organisationprofessional organizationlowland burrowing treefrogmary wollstonecraft godwininternal-combustion enginetechnology administrationedward goldenberg robinsonalgernon charles swinburnepaul johann ludwig von heysereaction-propulsion engineposterior meningeal arterymax karl ernst ludwig plancksensorineural hearing losselectromagnetic delay lineelectromagnetic intrusionelectromagnetic radiationinstrumental conditioningelizabeth barrett browningoffice of inspector generalgeorge washington goethalsarchitectural engineeringsir roger gilbert bannisterpleading in the alternativegastrointestinal disorderhudson bay collared lemmingsilver wedding anniversaryaccommodating lens implanttheory of organic evolutionmetacarpophalangeal jointgastrophryne carolinensisbell-type annealing furnaceblack and gold garden spiderelectronic data processingreal gross national product
...View all with 24 letters...
Phrases (43)
central intelligence agencyantonio allegri da correggiointernational organisationinternational organizationsir arthur stanley eddingtoncharles frederick menningerveronica anagallis-aquaticagreat plains of north americaorganic light-emitting diodecanadian arctic archipelagounited states virgin islandsjacques etienne montgolfieromnidirectional radio rangegasterophilus intestinalisbattle of pittsburgh landingcygnus columbianus bewickiiangus frank johnstone wilsonelectromagnetic transduceroffice of naval intelligenceproto-indo european languageislamic state of afghanistandistinguished conduct medalmargaret munnerlyn mitchellplastic blow molding machinedow-jones industrial averagecapital of the united kingdomvon recklinghausen's diseaseconsciousness-altering drugthyroid-stimulating hormonelaw of conservation of energynatural language processingcongenital afibrinogenemiapurple-staining cortinariusantennaria plantaginifoliadefense intelligence agency
...View all with 25 letters...
Phrases (36)
charlotte anna perkins gilmanfrances eliza hodgson burnettpolicing and enforcement costmalignant neoplastic diseasepersonal information managersir charles scott sherringtonbooker taliaferro washingtonevelyn arthur saint john waughassyrian neo-aramaic languagelouis antoine de bougainvillealpine enchanter's nightshadeentandrophragma cylindricumbenign prostatic hyperplasiaelectromagnetic interactiongallinago gallinago delicatacalifornia single-leaf pinyonalternating electric currentfinancial management serviceplastic blow moulding machinefollicle-stimulating hormonecommunications intelligencegeneralized anxiety disordermilitary intelligence agencybetty naomi goldstein friedanneohygrophorus angelesianuscalamintha nepeta glantulosathyrotropin-releasing factortissue plasminogen activatornarrow-leaved everlasting peacoptis trifolia groenlandicatrigonella ornithopodioideshousing and urban developmentsir frederick gowland hopkinsneuromuscular blocking agentblack september organization
...View all with 26 letters...
Phrases (27)
aleksandr sergeyevich pushkinaugustus welby northmore puginbeta-adrenergic blocking agentnuclear regulatory commissiontricyclic antidepressant drugthyrotropin-releasing hormonecryptobranchus alleganiensistopical prostaglandin eyedropapplication-oriented languagewrangell-st. elias national parkinhalation general anestheticnongovernmental organizationrepublic of trinidad and tobagoadeline virginia stephen woolfequivalent-binary-digit factorjohannes evangelista purkinjepleuropneumonialike organismmujahidin-e khalq organizationmartin luther king jr's birthdaywilliam iv of the united kingdomronald george wreyford norrisharchaeological planning frameunited states attorney generalnasopharyngeal leishmaniasisnegative reinforcing stimulussclerosing leukoencephalitiscount alessandro di cagliostroPhrases (33)
ephippiorhynchus senegalensisbattles of lexington and concordcapital of serbia and montenegrocriminal investigation commandfederal housing administrationludwig josef johan wittgensteinsergei mikhailovich eisensteinrevolutionary communist leagueimperial japanese morning glorysmall-for-gestational-age infanthypothalamic releasing hormoneinhalation general anaestheticcentral intelligence machinerygates of the arctic national parkkonstantin sergeevich alekseevcentral vein of suprarenal glandgiovanni virginio schiaparellifederal bureau of investigationmilitary intelligence section 5military intelligence section 6aboriginal australian languagebovine spongiform encephalitismelanocyte-stimulating hormoneinternational logistic supportgrowth hormone-releasing factorpan troglodytes schweinfurthiievangelical and reformed churchsatureja calamintha glandulosaalpha-adrenergic blocking agenthepatolenticular degenerationsergei vasilievich rachmaninovtime-delay measuring instrumentpreemployment training programPhrases (22)
male internal reproductive organsouth-central dravidian languagegiovanni pierluigi da palestrinascrutin uninominal voting systemcongregational christian churchreceived pronunciation languagehydrangea macrophylla hortensisnational intelligence communityfrancisco jose de goya y lucientesdirector of central intelligenceelectron paramagnetic resonancecombined transport bill of ladinggeneral services administrationgrigori aleksandrovich potemkinhealth maintenance organizationvigna unguiculata sesquipedalisbergen-belsen concentration campgaius plinius caecilius secundusenvironmental protection agencyage-related macular degenerationsergei vasilievich rachmaninoffimaginary part of a complex numberPhrases (16)
glutamic oxalacetic transaminasedepartment of energy intelligencerepublic of bosnia and herzegovinauniversal gravitational constantaugust friedrich leopold weismannnontricyclic antidepressant drughenri rene albert guy de maupassant1st viscount montgomery of alameinbachelor of science in engineeringinternational labor organizationfood and agriculture organizationferdinand victor eugene delacroixmarital communications privilegegestalt principle of organizationguadalupe mountains national parkunited states intelligence agencyPhrases (26)
great smoky mountains national parkbureau of intelligence and researchdigital communications technologysidonie-gabrielle claudine colettepalestine liberation organizationpalestinian popular struggle frontelectromotive drug administrationislamic great eastern raiders-frontgeorges eugene benjamin clemenceaufemale internal reproductive organglutamic oxaloacetic transaminasewernher magnus maximilian von braunmaria luigi carlo zenobio cherubiniinternational atomic energy agencymarie dolores eliza rosanna gilbertinternational intelligence agencyinternational labour organizationmarine corps intelligence activityovulation method of family planningevangelical united brethren churchluigi barnaba gregorio chiaramontinorth atlantic treaty organizationworld meteorological organizationgreat eastern islamic raiders' frontvocational rehabilitation programmary godwin wollstonecraft shelleyPhrases (13)
african american vernacular englishpeople against gangsterism and drugsweakly interacting massive particlestraight-line method of depreciationrevolutionary justice organizationobject-oriented programing languagecount nikolaus ludwig von zinzendorfnonsteroidal anti-inflammatory drugedmond louis antoine huot de goncourtyevgeni aleksandrovich yevtushenkogiovanni jacopo casanova de seingaltattorney general of the united statesunited self-defense group of colombiaPhrases (16)
linear-quadratic-gaussian controllermusical instrument digital interfaceinternational maritime organizationstandard generalized markup languagecongress of industrial organizationsobject-oriented programming languageelizabeth cleghorn stevenson gaskellprivilege against self incriminationgeneral agreement on tariffs and tradekonstantin sergeyevich stanislavskymercury-in-glass clinical thermometerfloating-point representation systeminternational law enforcement agencysubacute sclerosing panencephalitisunited states intelligence communityphysiological jaundice of the newbornPhrases (17)
relational database management systemfederal national mortgage associationnero claudius caesar drusus germanicusinterstitial cell-stimulating hormonefriedrich august kekule von stradonitzeuropean law enforcement organisationvena centralis glandulae suprarenalisfunctional magnetic resonance imaginghermann ludwig ferdinand von helmholtzfour-stroke internal-combustion enginemassachusetts institute of technologyfederal home loan mortgage corporationforeign intelligence surveillance actbaron hugh caswall tremenheere dowdingeinstein's general theory of relativitycomputerized axial tomography scannergreat proletarian cultural revolutionPhrases (11)
revolutionary organization 17 novembercanadian security intelligence serviceimmaculate conception of the virgin marycockcroft and walton voltage multipliernational socialist german workers' partyimmigration and naturalization serviceanti-imperialist international brigadefederal law enforcement training centerballistic missile defense organizationsubacute sclerosing leukoencephalitisfraxinus pennsylvanica subintegerrimaPhrases (8)
jakob ludwig felix mendelssohn-bartholdyforeign intelligence surveillance courtnational geospatial-intelligence agencyalbert francis charles augustus emmanuelcriminal intelligence services of canadanatural language processing applicationtiberius claudius drusus nero germanicusdeclining balance method of depreciationPhrases (7)
million floating point operations per secondbillion floating point operations per seconddefense reutilization and marketing servicedirectorate for inter-services intelligencegates of the arctic national park and preservebaron hermann ludwig ferdinand von helmholtzenzyme-linked-immunosorbent serologic assay